1. Plants shrubs, trees, or lianas; embryo with differentiated
cotyledons. . . . . . .4
4. Pollen shed in tetrads; seeds lacking a vascular bundle in raphe;
anthers usually lacking a fibrous endothecium (except Bejaria),
opening by pores or slits . . . .5
5. Flowers wind-pollinated, inconspicuous, imperfect, with perianth of
distinct tepals, the stigma flabellate to pinnatifid; leaves always
ericoid . .
. . . .Ericoideae, Empetreae.
5. Flowers usually ± showy and insect or bird pollinated, usually
perfect, with perianth of a calyx and usually sympetalous corolla,
the stigma truncate to ± capitate, rarely cup-shaped or
funnel-shaped ;leaves ericoid to broad and flat. .6
6. Anthers tetrasporangiate, dithecal; leaves pinnate to palmately
veined;stamens usually twice the number of corolla lobes,
occasionally reduced and equaling the corolla lobes . . . . . . . 13
13. Ovary superior; anthers usually lacking tubules. . . . . . . 14
14. Anthers inverting early in development of the flower; fruit
lacking an endocarp, usually dehiscent; ± several to numerous
ovules per locule. . . . . . . . . . . . . . . . . . . . . . 15
15. Corolla persistent; leaves ericoid. . . Ericoideae, Ericeae,
p.p.
15. Corolla deciduous; leaves ericoid to broad and flat . . . 16
16. Fruit a ± septicidal capsule . . . . . . . . . . . . . . 17
17. Stamens epipetalous 2/3 length of filament. . . Diplarche
(Rhodoreae)
17. Stamens not adnate to corolla. . . . . . . . . . . . . 18
18. Endothecium present and fibrous; flowers more than 5-
merous. . . . . . . . . . . . Bejarieae, (Bejaria)
18. Endothecium lacking, or present and lacking a fibrous
layer; flowers 4 or 5-merous. . . . . . . . . . . .19
19. Seeds papillose; pedicel lacking bracteoles;
filaments smooth. Daboecia (Ericoideae, Ericeae).
19. Seeds not papillose; pedicel with 2 bracteoles;
filaments roughened or unicellular-pubescent. .20
20. Leaves ericoid; viscin threads usually absent
. . . . . . . . . . . . . . . . . . . . . . 21
21. Petals distinct. Ericoideae, Bejarieae
21. Petals connate. .Phyllodoce (Ericoideae,
Phyllodoceae)
20. Leaves not ericoid, although sometimes with
revolute margins; often with viscin threads
. . . . . . . . . . . . . . . . . . . . . 22
22. Pedicel usually not articulated with
flower; flowers usually zygomorphic and
with spots or a blotch; bracts usually
scarious; ovary ± ovoid to cylindric;
perulae usually present. . . . Ericoideae,
. . . . . . . . . . . . . . Rhodoreae p.p.
22. Pedicel articulated with pedicel; flowers
usually actinomorphic, lacking spots or a
blotch; bracts usually not scarious; ovary ±
globose to subglobose; perulae lacking. . .
. . . . . . . Ericoideae, Phyllodoceae p.p.
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