6. Anthers tetrasporangiate, dithecal; leaves pinnate to palmately
veined;stamens usually twice the number of corolla lobes,
occasionally reduced and equaling the corolla lobes . . . . . . . 13
13. Ovary inferior; anthers with tubules . Vaccinioideae, Vaccinieae
13. Ovary superior; anthers usually lacking tubules. . . . . . . 14
14. Anthers inverting early in development of the flower; fruit
lacking an endocarp, usually dehiscent; ± several to numerous
ovules per locule. . . . . . . . . . . . . . . . . . . . . . 15
15. Corolla deciduous; leaves ericoid to broad and flat . . . 16
16. Fruit usually a loculicidal capsule, occasionally a berry.23
23. Leaves not ericoid, although they may have a revolute
margin; bud scales present; anthers usually with
disintegration tissue; fibers associated with vascular
tissue in petiole and leaf midrib. . . . . . . . . . .25
25. Secondary phloem with bands of fibers; stamens
lacking appendages or with 2 spurs at anther-filament
junction or on upper portion of filament; leaf
epidermis ± lignified; filaments usually S-shaped. .
. . . . . . . . . . . . . . . Vaccinioideae,Lyonieae
25. Secondary phloem lacking bands of fibers (except
Zenobia, which has 4 awns on anthers); stamens
lacking appendages or with 2 or 4 awns arising ± from
anther apex; leaf epidermis not lignified (except
Andromeda, which has 2 awns on anthers); filaments
straight. . . . . . . . . . . . . . . . . . . . . 26
26. Leaves lacking multicellular hairs, except for ±
globose, non-glandular hairs associated with
serrations in Zenobia; stomata anomocytic. . .
. . . . . . . . . . Vaccinioideae, Andromedeae
26. Leaves with scattered gland-headed, multi-
cellular hairs; stomata usually paracytic. . 27
27. Pedicel not articulated; inflorescence
terminal; flowers borne on shoots of
the current season; anthers not appendaged;
fruits ± elongate-ovoid. . . .
. . . . . . . . Vaccinioideae, Oxydendreae
27. Pedicel articulated with flower;
inflorescences usually axillary; flowers
usually on shoots of the previous season;
anthers often with 4 "apical" appendages (awns); fruits
± globose to subglobose. . . .
. . . . . . . . .Vaccinioideae, Gaultherieae |